The animals were euthanised following 'complications' during an operation on the alpha male wolf
The animals were euthanised following "complications" during an operation on the alpha male wolf, Loki, at Camperdown Wildlife Centre in Dundee. Despite Loki's initially successful operation, there were "subsequent complications" - and park bosses took the "difficult decision" to save the animal "from further pain" on Monday.
In a post on Facebook, the centre said: "We are devastated to announce that our alpha male wolf, Loki, has been humanely euthanised today. "The other four wolves in the pack have been exhibiting unusually anxious and abnormal behaviour since the operation, and it is with great sadness that these wolves have also been euthanised as a result.
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