It follows two crashes involving the aircraft in less than five months in which 346 have died.
Boeing said the software enhancements included limiting stabiliser trim commands in response to erroneous readings, pilot displays, operation manuals and crew training.
Authorities in China, South Africa, Singapore, Indonesia and Mexican airline Aeromexico have followed suit - but several airlines serving UK airports, including TUI and Norwegian, are continuing to fly the planes. Colleagues at the UN fisheries and aquaculture department described Ms Toole as a"wonderful human being", while her father said she was a"very soft and loving" woman.
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